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Layed timing of peduncle elongation may well afford wind and frost protection throughout anthesis, when later enhancing fruit dispersal. Peduncle elongation does not normally accompany fruiting or might be minimal, but even in specimens with many subsessile fruiting capitula nonetheless embedded inside the leaf mat, occasional remains of significantly longer peduncles may be found (e.g. Barbour 3427, Sagastegui 12242). Notably, the only specimen from Peru and Ecuador with peduncles already elongate (two cm) at anthesis was also the only specimen reportedly collected from “montane forest,” where it was stated to be rare (Cano 16840). This plant may perhaps represent a taxonomically distinct variant, but for now is treated as an environmental kind. In Colombian populations in the Eastern Cordillera, early peduncle elongation could be more frequent. Other characters MedChemExpress SPDB displaying wide variation in Peru and Ecuador are involucral bract length, capitulum size, and flower and seed size. By way of example, the type of P. loxensis (Loja) contrasts sharply with that of P. espinosianus (Azuay) by its much smaller involucral bracts, capitula, flowers, and seeds. Indeed, in most plants from Azuay, in the Cerro Fierro-Urcu (Loja), and in some localities in Peru (Ja , Bagua, Celend ), outer involucral bracts are ca.2.six.0 mm, together with the tips typically surpassing the significant capitula four mm in diameter (cf. Fig. 2H). In contrast, most specimens from southern Loja, the western Cordillera of Peru, and southern Amazonas, Peru, have involucral bracts about 1.6.6 mm long and capitula 3 mm in diameter (Fig. 2I ). The length on the pistillate flower sepals was measured in 18 specimens of Peru and Ecuador and was imperfectly correlated with capitulum size. Compact flowers (sepals 1.four.8 mm) were found within the small-capitate plants from southern Loja and northwestern Peru, and a few large-capitate plants of Azuay; all other specimens had been large-flowered (sepals ca. 1.8.4 mm long). Seed size showed the strongest geographic correlation, together with the smallest seeds observed inside the small-flowered specimens from southern Loja and also the western cordillera of Peru (0.53.63 mm, 5 samples), when elsewhere seeds were bigger (0.67.8 mm, eight samples). Among the large-flowered collections, these of Amazonas, Peru, south of the mouth from the Rio Utcubamba (ca. 6S latitude) are distinguished by the unusually broad female petals (l/w ratio 1.8.9 vs. three.5.0 elsewhere), broadly rounded to even truncate-emarginate at apex (Fig. 12I), and by an ambiguous pattern of sepal thickening, each characters suggesting intermediacy with P. pilosus var. leoniae, of San Mart . Also shared with that selection would be the funnelform nectaries (Fig. 12I) with unusually massive, spreading papillae (also identified in Sagastegui 12242, Celend ), in comparison for the commonly penicillate nectaries with tiny papillae identified elsewhere. Paepalanthus pilosus PubMed ID:http://www.ncbi.nlm.nih.gov/pubmed/20107869 in Colombia needs additional careful study. In the Eastern Cordillera, especially in disturbed paramo close to Bogot you will find a lot more pronounced morphological extremes inside the species than in other components of its distribution. Right here, leaf lengths range up to 3 cm, capitula from 3.5 mm in diameter, peduncles are occa-Nancy Hensold / PhytoKeys 64: 17 (2016)Figure 12. Pistillate flowers and fruits of P. pilosus varieties. A-K P. pilosus var. pilosus (A-B J gensen 2366 C-F Cano 16840 G Cuatrecasas 25574 H MacDougal 4463 I Wurdack 1616 J lgaard 9557 K Boeke 2133) A-B Flower in anthesis with detail of gynoecial nectaries C Flower in anthesis D Gynoecium E.

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